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Webster 1913 Edition
Rhachis
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1. 
(Anat.) 
The spine. 
2. 
(Bot.) 
(a) 
The continued stem or midrib of a pinnately compound leaf, as in a rose leaf or a fern. 
(b) 
The principal axis in a raceme, spike, panicle, or corymb. 
3. 
(Zool.) 
(a) 
The shaft of a feather. The rhachis of the after-shaft, or plumule, is called the hyporhachis. 
(b) 
The central cord in the stem of a crinoid. 
(c) 
The median part of the radula of a mollusk. 
(d) 
A central cord of the ovary of nematodes.